
The first step in matching patterns in nature to evolutionary process models is to determine what the expected pattern from each model. The biggest problem in matching species complex patterns to process models is that none of the three monophyletic processes are mutually exclusive. In any one monophyletic species complex, you might find any combination of these three processes at different times or between different populations. The result will blur the edges between the expected patterns. Pigeon-holing a pattern into one of these three models without very clear evidence would be grossly reductionistic in respect to the complexity of evolutionary process. It is more conservative to simply determine if the complex is monophyletic or not. Any further hypothesis concerning the causative evolutionary processes can only be tenative and, unless there is a clearcut pattern, may not be definitively testable.